In 1965, this actress played the first officer of a ship in a rejected pilot episode, partially rejected because NBC was convinced no one would accept a woman in a strong commanding role. When the series was later picked back up, this actress played the ship's nurse. In a much later spin-off series, this actress played a comedic over-domineering mother, though in later episodes she was shown to be complex and strong, capable of meaningful relationships with a David Ogden Stiers played scientist forced to commit suicide, and with an embarrassed shapeshifter. Identify this actress who sometimes hyphenated her name with that of her husband, Gene Roddenberry.
Majel Barrett